Taxonomy of Opiliones - Minor authors

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Assorted papers by various authors

Please note that the expression "minor authors" does not express any judgement of value. It refers only to authors who focused on other things and therefore produced very few papers on Opiliones. A "minor author" for Opiliones may be a "major author" for spiders for example.


  • BAS (contributed PDF Merrett 1984)
  • Christine Rollard & Mark Judson (allowed access to facilities in MNHN, Paris, 2013 - PDFs Falconer 1927a, Gasperini 1891-1892).
  • Jan Beccaloni & Sarah Sworder (allowed access to facilities in the NHM, London, 2013 - Bristowe 1928).
  • Laura Leibensperger (provided reprint of Katayama & Post 1974).
  • MNRJ library (provided reprint of Lotz 2002).Pavel Bezděčka (contributed PDF of Dolansky 1997)
  • Janet Beccaloni + Milena Kury (reavers of London libraries, provided photocopies of Bacelar ; 1950; Ermolajev 1937)
  • Theo Blick (contributed PDFs - Herbst, 1798, 1799)
  • Luca Cavigioli (contributed PDFs Carr 1919, Olsen 1995).
  • Emmanuel Delfosse (contributed PDF - Berland 1949)
  • Jason Dunlop, Anja Friederichs, Ian S. Kury & Hans-Ulrich Raake (participated in my biblio-hunt in ZMB, Berlin 2009 -- PDFs Müllers)
  • Ian S. Kury (contributed PDFs - Delany 1959; Dugès 1885; Walker 1928; provided photocopies - Janczyk 1956, Roth 1851)
  • Milena Kury (contributed Graaf 1882)
  • Gonzalo Giribet (contributed PDFs - Silvestri 1904)
  • Lars Friman (contributed PDF - Costa 1885)
  • Dmitri Logunov (provided photocopies - Farzalieva & Esyunin 2000)
  • Plamen Mitov (contributed PDF - Wang 1941).
  • John Ponting (indicated link to Stolicka 1869, contributed PDFs Baert et al. 1995; Bowser 2008; Bragg & Holmberg 2009; Cantrell 1980; Gertsch 1939; Moritz 1973)
  • Nataly Snegovaya (contributed PDF - Bogachev 1951).
  • Axel Schönhofer (contributed PDF - Müller & Schenkel 1895)
  • Daniele Souza (digitalized - Cawley 1995; Dugès 1886).
  • Wojciech Staręga (contributed PDFs - Lotz 2009, 2011).
  • Andras Varga (contributed PDF - Bokor 1996).
  • Chao Zhang (contributed PDF - Wang 1953).


DISCLAIMER — This resource was first intended as for private use of the members of the arachnology lab of Museu Nacional, but later we thought "why not to share this with the world?". Eventually if greedy lawyers (redundance...) start to bother us with copyright matters, etc, we may have to be forced to quit the project and keep this just to ourselves.
Abbott, R.H.R. (1981) A new opilionid to Great Britain. Newsletter of the British Arachnological Society, 30, 4.
xrx na fila scan Arnold, R.A., (1993) Harvestmen, pp. 537–539. In: Endangered wildlife of the world. Vol. 4, FOD-IBI, Marshall Cavendish, New York, pp. 433–576.
paper needed Askew, R.R. (1976) Dicranopalpus caudatus Dresco (Arachnida: Opiliones) in South Wales. The Entomologist's monthly Magazine, 111 [“1975”], 100.
Babić, K. (1916) Opilionidi hrvatskog zemaljskog zoološkog muzeja u Zagrebu [Die Opilioniden des kroatischen zoologischen Landesmuseums zu Zagreb]. Glasnik hrvatskoga prirodoslovnoga društva, Zagreb, 28, 169–179.
Bacelar, A. (1928) Aracnídios Portuguêses. III. Catalogo sistematico dos Aracnidios de Portugal citados por diversos Autores (1831–1926). Bulletin de la Société Portugaise des Sciences Naturelles, Lisbonne, 10(17), 169–203. [Issued 28 March 1928].
Bacelar, A. (1950) Notas acerca dos aracnídeos do ultramar português. Colóquios Realizados na Junta de Investigações Coloniais de 11 de Novembro de 1949 a 23 de Junho de 1950, Lisboa, 17, pp. 1–23 + pl. I-III.
Baert, L., Maelfait, J.-P. & Desender, K. (1995) Distribution of the arachnid species of the orders Scorpiones, Solifugae, Amblypygi, Schizomida, Opiliones and Pseudoscorpiones in Galápagos. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique. Entomologie, 65, 5–19.
  Bailey, G.W.D. (1973) Irish spiders and harvestmen. Irish Naturalists' Journal, 17(12), 421.
Basset, Y. (1985) Les peuplements d'arthropodes sur Pinus mugo Turra dans les tourbières du Haut-Jura Neuchâtelois. Bulletin de la Société Neuchâteloise des Sciences Naturelles, (4) 108, 63–76.
  Bauer, Christian & Prieto, Carlos E. (2009) Three new Assamiidae (Arachnida: Opiliones) from Cameroon, with a redescription of Chilon robustus and comments on related species. Zootaxa, 2059, 1–22.
Bell, R.T. (1974) A European harvestman in North America (Phalangida, Phalangiidae). Entomological News, 85(5-6), 154.
  Beretzk, P., Csongor, G., Horvath, A., Karpati, A., Kolosvary, G., Szabados, M., Szekely, M. (1957) Das Leben der Tisza. I. Uber die Tierwelt der Tisza und ihrer Inundationsgebiete. Acta Biologica, Szeged, nova series, 3, 81–108.
Berland, L., 1949. Ordre des Opilions. In: P. Grassé, Traité de Zoologie, Tome 6. Onychophores, tardigrades, arthropodes, trilobitomorphes, chélicérates. Masson, Paris, pp. 761–793, figs 553–597.
Birstejn Ja. A. 1950. Fauna of caves of the Western Caucasus. Journ. of Zoology, Moscow, 29: 354–366 [in Russian].
Bivort, B.L. de & Giribet, G. (2004) A new genus and species of cyphophthalmid from the Iberian Peninsula with a phylogenetic analysis of the Sironidae (Arachnida, Opiliones, Cyphophthalmi). Invertebrate Systematics, 18, 7–52.
  Bivort, B.L. de & Giribet, G. (2010) A systematic revision of the South African Pettalidae (Arachnida: Opiliones: Cyphophthalmi) based on a combined analysis of discrete and continuous morphological characters with the description of seven new species. Invertebrate Systematics, 24(4), 371-406.
Bivort B.L. de, Clouse R.M. & Giribet G. (2010) A morphometrics-based phylogeny of the temperate Gondwanan mite harvestmen (Opiliones, Cyphophthalmi, Pettalidae). Journal of Zoological Systematics and Evolutionary Research, 48(4), 294-309.
Bivort B.L. de, Clouse R.M. & Giribet G. (2012) A cladistic reconstruction of the ancestral mite harvestman (Arachnida, Opiliones, Cyphophthalmi): portrait of a Paleozoic detritivore. Cladistics, 28, 582–597.
Bogachev, A.A. (1951) Отряд сенокосцы [Otriad senokostsi; Order Opiliones]. In: Alizade, A.N. et al. (Eds.), Животный мир Азербайджана [Zhivotniy mir Azerbaidzhana; Animal World of Azerbaijan], Baku, AS AzSSR, Inst. of Zoology, pp. 405–406 (in Russian).
Bokor, Z. (1996) New occurrence of Astrobunus laevipes (Canestrini, 1872) (Arachnoidea, Opiliones, Phalangiidae) in Hungary (in the valley of river Rakaca — Cserehát). Acta Biologica, Szeged, 41, 89–91.
reprint na fila Bokor, Z. (1998) Data on the Opiliones (Arachnida) fauna of the Rakaca valley (Cserehát, Hungary). Folia Entomologica Hungarica, 59, 221-225.
Bowser, Matthew (2008) Notes on a Kenai Sabacon (Opiliones: Sabaconidae). Newsletter of the Alaska Entomological Society, 2(1), 5-7. [Issued February 2008].
  Bragg, P.D. & Leech, R.E. (1972)  Additional records of spiders (Araneida) and harvestmen (Phalangida) for British Columbia. Journal of the Entomological Society of British Columbia, 69, 67–71.
PDF from JOA local copy. Bragg, P. D. & Robert G. Holmberg, 1974. Platybunus triangularis and Paroligolophus agrestis: two phalangids introduced to North America (Arachnida, Opiliones). The Journal of Arachnology, 2(2): 127.
Bragg, P.D. & Holmberg, Robert G. (2009). The Harvestmen (Arachnida, Opiliones) of British Columbia. Journal of the Entomological Society of British Columbia, Vancouver, 106, 29–37. [Issued Dec. 2009].
Bristowe, W.S. (1928) The Spider Fauna of the Western Islands of Scotland. The Scottish naturalist, 165 [“1926”], 88–94.
Cantrell, B. K., 1980. Additional Australian harvestmen (Arachnida: Opiliones). Journal of the Australian Entomological Society, Brisbane, 19: 241-253.
Carr, L.A. (1919) The spiders, harvestmen and pseudoscorpions of Lichfield and neighborhood. Transactions North Staffordshire Field Club, 53 1918–19, 71–87.
Cawley, M. (1995) Dicranopalpus ramosus (Simon) (Arachnida: Opiliones), new to Ireland. Irish Naturalists' Journal, 25(4), 153.
  Cawley, M. (1999) New Irish vice-county records for harvestspiders (Arachnida: Opiliones). Bulletin of the Irish Biogeographical Society, 23, 82–88. Published:  15 December 1999.
  Cawley, M. (2002) A review of the Irish harvestmen (Arachnida: Opiliones). Bulletin of the Irish Biogeographical Society, Dublin, 26, 106–137. [Issued 15 December 2002].
  Celiński, Florian (1955) Projektowane rezerwaty leśne w Puszczy Bukowej pod Szczecinem [The proposed forest reserves in the Beech Forest near Szczecin]. Chrónmy przyrod¸e ojczyst¸a : organ Pa´nstwowej Rady Ochrony Przyrody [= Instytut Ochrony Przyrody Polskiej Akademii Nauk]. Ojez., 11(4), pp.3–15, ff. 1–12. Krakow.
  Cerruti, Marcello (1968) Materiali per un primo elenco degli artropodi speleobii della Sardegna. (In memoria di Saverio Patrizi). Fragmenta Entomologica, Roma, 5(3) [“1967”], 207–257.
external link Colenso, W., 1882. On some newly-discovered New Zealand arachnids. Transactions and Proceedings of the New Zealand Institute, 15: 165-173.
Costa, Achille (1885) Notizie ed osservazioni sulla geo-fauna sarda. Memoria quarta. Atti della Reale Accademia delle scienze fisiche e matematiche di Napoli, (serie seconda) 1(13) [“1888”], 1–31.
  Cuppen, J.G.M. (1994) Dicranopalpus ramosus, a new species of harvestman for The Netherlands (Opilionida: Phalangiidae). Entomologische Berichten, Amsterdam, 54(9), 176–178.
  Curcic, Bozidar P.M., Ivo M. Karaman & V.T. Tomic, 1999. On some new and little known epigean harvestman (Opiliones, Arachnida) from West Serbia, Yugoslavia. Archives of biological sciences, Belgrade, 51(4): 61-62.
Cuvier, G. (1795) Description de deux espéces nouvelles d'Insectes. Le Faucheux a 4–dentelures. Magazin Encyclopédique, (N.S.), 1(2), 205–207, plate 2.
Dalmas R., Le Comte de (1922) Catalogue des araignées récoltées par le Marquis G. Doria dans l'île Giglio (Archipel toscan). Annali del Museo Civico di Storia Naturale Giacomo Doria, Genova, Series 3, 10(50), 79–96.
  Dancau, D. & Tabacaru, I. (1964) Observatii zoogeografice asupra faunei cavernicole din Oltenia si Banat. Lucrarile Institutului de Speologie «Emil Racoviţa», 3, 293–340 figs. 1–9. [In Romanian].
Delany, M.J. (1959) Harvestmen (Arachnida, Opiliones) from the Island of Colonsay, Inner Hebrides. The Entomologist's Monthly Magazine, 95, 158.
Derkarabetian, S. & Hedin, M. (2014) Integrative taxonomy and species delimitation in harvestmen: a revision of the Western north american genus Sclerobunus (Opiliones: Laniatores: Travunioidea) PloS One 2014 9(8): e104982
Derkarabetian S., Ledford, J. & Hedin, M. (2011) Genetic diversification without obvious genitalic morphological divergence in harvestmen (Opiliones, Laniatores, Sclerobunus robustus) from montane sky islands of western North America. Molecular Phylogenetics and Evolution, 61, 844-853.
Derkarabetian, S, DB Steinmann & M Hedin (2010) Repeated and time-correlated morphological convergence in cave-dwelling harvestmen (Opiliones, Laniatores) from montane western North America. PLOS One, 5(5), e10388, 1–13.
Dolanský, Jan (1997) Nález klepítníka cleneného Ischyropsalis hellwigi (Panzer, 1794) - (Opilionida) u Novych Hradu (Vychodní cást okresu Chrudim). [Find of Harvestman Ischyropsalis hellwigi (Panzer, 1794) (Opilionida) near Nové Hrady (east part of Chrudim district)]. Východočeský Sborník Přírodovědný,Práce a Studie, 5, 187–188 (in Czech).
Dugès, Alfredo (1885) Opilio ischionotatus, Alfredo Dugès. Segador de ancas manchadas de blanco. La Naturaleza, Periódico científico del Museo N. de Historia Natural y de la Sociedad Mexicana de Historia Natural, México 7, 194–196 + plate 2a.
Dugès, Alfredo (1886) Je trouve décrit (Soc. entom. Belge, Compte rendu, février 1886) un Egaenus mexicanus par M. L. Becker. Annales de la Société Entomologique de Belgique, Bruxelles, 30: cxxvi.
Ermolajev, W. (1937) Beiträge zur Spinnentier-Fauna der Stadt Tobolsk (West-Sibirien). Festschrift zum 60. Geburtstage von Professor Dr. Embrik Strand, Rigā, 2, 519–524.
Falconer, W. (1927a) A Contribution to the arachnid fauna of Mid-Wales. The Naturalist,London,840, 27–29.
Farzalieva, G.Sh. & Esyunin, S.L. (2000) The harvestman fauna of the Urals, Russia, with a key to the Ural species (Arachnida: Opiliones). Arthropoda Selecta, 8(3) [1999], 183–199.
Gasperini, R. (1891) Prilog fauni dalmatinskih Pauka (Araneae et Opiliones) [Contributions to the fauna of Dalmatian Arachnida]. Godisnij izvejestaj vel. Realke Splitu, 1890-91, 1–18 (in Croatian).
Gasperini, R. (1892) Prilog k dalmatinskoj fauni (Isopoda. – Myriopoda. – Arachnida). Godisnij izvejestaj vel. Realke Splitu, 1891/92, 1–22 (in Croatian).
Geer, Charles de, 1778. Des Faucheurs (pp 163–175, plate 10). In: Mémoires pour servir à l'histoire des insectes. Tome 7. Pierre Hesselberg, Stockholm, xii + 950 pages + 49 plates.
Gertsch, W.J. (1939) Report on a collection of Arachnida from the Chisos Mountains. Contributions of the Baylor University Museum, 24, 17–26.
Graaf, Henri William de (1882) Sur la construction des organes génitaux des Phalangiens. E. J. Brill, Leiden, 95 + 31 pp, 35 pl. SEPARATE PDF WITH PLATES
Havranek, L. & Molnár, H. (1965) Preliminary report on the Arachnoidea-fauna of the Tisza-valley. Tiscia, 1, 93–107.
Helversen, O. von & Martens, J. (1972) Unrichtige Fundort-Angaben in der Arachniden-Sammlung Roewer. Senckenbergiana biologica, 53(1/2), 109–123.
Herbst, J.F.W. (1798) Naturgeschichte der Insecten-Gattung Opilio. In: Herbst, J.F.W. Natursystem der ungeflügelten Insekten, vol. 2 [of 4]. Gottlieb August Lange, Berlin, iv + 26 pp., 5 plates.
Herbst, J.F.W. (1799) Forsetzung der Naturgeschichte der Insectengattung Opilio. In: Herbst, J.F.W., Natursystem der ungeflügelten Insekten, vol. 3 [of 4]. Gottlieb August Lange, Berlin, iv + 30 pp., plates 6–10.
Janczyk, F.S.W. (1956) Anatomie von Siro duricorius Joseph im Vergleich mit anderen Opilioniden. Sitzungsberichte der Österreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien, Abteilung I, Biologische Wissenschaften und Erdwissenschaften, 165, 475–522, figs 1–28.
Katayama, R.W. & Post, R.L. (1974) Phalangida of North Dakota. North Dakota Insects, 9, l–40.
Le Roi, O. (1914) Zur Kenntnis der Opilioniden-fauna von Norddeutschland. Schriften der Physikalisch-Ökonomischen Gesellschaft zu Königsberg in Preußen, Königsberg, 54, 257–272.
Li, Shu-qiang & Song, Da-xiang (1993) A revision of Chinese Harvestmen, with a note on their geographical distribution. Scientific Treatise on Systematic and Evolutionary Zoology, 2, 237-245.
Lotz, L.N. (1999) Distribution of the Opilionid Polycoryphus asper Loman, 1902 in Southern Africa. Newsletter of the British Arachnological Society, 86, 13–14.
Lotz, L.N. (2002) The Opiliones (Arachnida) of the Free State Province, South Africa. Navorsinge van die Nasionale Museum, Bloemfontein, 18(8), 161-188. [Published: November 2002].
Lotz, Leon (2009) Harvestmen (Arachnida: Opiliones) in southern Africa - an annotated catalogue with notes on distribution. Navorsinge van die Nasionale Museum, Bloemfontein,, 25(1), 1-45.
Lotz, L.N. (2011) Three new harvestmen species from southern Africa (Arachnida: Opiliones: Caddidae, Neopilionidae, Assamiidae). Journal of Afrotropical Zoology, 7: 3–8.
PDF from JOA local copy. McGhee, Charles Robert, 1977 (“1975”). The politum group (bulbate species) of Leiobunum (Arachnida: Phalangida: Phalangiidae) of North America. The Journal of Arachnology, 3(3): 151-163.
PDF from JOA local copy.

McGhee, Charles Robert, 1978 (“1977”). Observations on the use of measurements in the systematic study of Leiobunum (Arachnida: Phalangida). The Journal of Arachnology, 5(2): 169-178.
Merrett, C.M. (1984) A further discovery of the harvestman Sabacon in Glamorgan. Newsletter of the British Arachnological Society, 40, 2.
Moritz, M. (1971) Die Typen der Arachniden-Sammlung der zoologischen Museums Berlin I. Opiliones. Mitteilungen aus dem Zoologischen Museum in Berlin, 47(1), 189–214.
Moritz, M. (1973) Neue und seltene Spinnen (Araneae) und Weberknechte (Opiliones) aus der DDR. Deutsche Entomologische Zeitschrift, N.Ser., Berlin, 20(1/3), 173–210.
Müller, F. & Schenkel, E. (1895) Verzeichnis der Spinnen von Basel und Umgegend, mit gelegentlicher Berücksichtigung ausserbaslerischer Schweizerarten. Verhandlungen der Naturforschenden Gesellschaft in Basel, Basel, 10, 691–824, pl. 13–14.
Müller, Hans-Georg (1983) Opilioniden-Funde aus der Schweiz und Frankreich (Arachnida: Opiliones). Entomologische Zeitschrift,  93(21) 315–318.
Müller, Hans-Georg (1989) Nemastoma dentigerum CANESTRINI 1873 aus der Wetterau, ein in Deutschland seltener Weberknecht (Arachnida: Opilionidat Nemastomidae). Beiträge zur Naturkunde der Wetterau, 9(1), 95–97.
Müller, Leon (1962) Les Phalangides dans le Grand-Duché de Luxembourg. Archives Institut Grand-Ducal Luxembourg, Section des Sciences naturelles, physiques et mathématiques, (N.S.) 28 [“1961”], 233-248.
Murphree, C.S., 1988. Morphology of the dorsal integument of ten opilionid species (Arachnida, Opiliones). The Journal of Arachnology, 16(2), 237–252.
Olsen, K.M. (1995) Opilio parietinus (De Geer, 1778) (Arachnida, Opiliones) does belong to the Norwegian fauna. Fauna norvegica B 42, 66–67.
Pack-Beresford, D.R. (1929) A revision of the Irish spider list. pp. 41–52. Report on recent additions to the Irish fauna and flora (Terrestrial and freshwater). Edited by R. Lloyd Praeger. Proceedings of the Royal Irish Academy, Dublin, (Section B. Biological, geological and chemical science) 39(1), 1–94, pl. 1.
Phillipps, W.J. & R.E.R. Grimmett (1932) Some new Opiliones from New Zealand. Proceedings of the Zoological Society of London, 1932(34), 731–740, 6 figs.
Roth, Johann Rudolph (1851) Ueber fossile Spinnen des litografischen Schiefers. Gelehrte Anzeigen d. Königlich Bayerische Akademie der Wissenschaften, München, 32, 164–167. [Issued 4 Feb. 1851].
Silvestri, F. (1904) Note arachnologiche II. Descrizione di un nuovo genere di Opilionidi del Chile. Redia: Giornale di Entomologia, Firenze, 2, 254–256 + pl. 23.
Silvestri, F. (1906) Descrizione di un nuovo genere di Opilionidi del Chile. Revista chilena de historia natural, Santiago, 10(1), 32–34.
Stoliczka, F. (1869) Contribution towards the knowledge of Indian Arachnoidea. Journal of the Asiatic Society of Bengal, Calcutta, 38(2), 201–251, plates 18–20.
Thor, Sig, 1933. Melloinia nom.nov. für Canestrinia (non A. Berlese 1881) M. Mello-Leitão 1931. Zoologischer Anzeiger, Leipzig, 103(7-8): 224.
Underwood, L.M. (1885) A Preliminary List of the Arthrogastra of North America (excluding Mexico). The Canadian Entomologist, 17(9), 162-169.
PDF from JOA local copy. Vázquez, Ignacio M. & James C. Cokendolpher, 1997. Guerrobunus vallensis, a new species of harvestman (Opiliones: Phalangodidae), from a cave in Valle de Bravo, State of México, México. The Journal of Arachnology, 25(3): 257-261.
Walker, Mary Elizabeth (1928) Revision of the order Phalangida of Ohio. Ohio State University Studies, Biological Survey Bulletin 19 (4), 153–175.
Wang, Feng-Chen (1941) Chinesische Spinnentiere. I. Opiliones. Zoologischer Anzeiger 135(5/6), 97–115. [Issued 15 August 1941].
Wang, Feng-Chen (1953) A list of Chinese spiders of the order Opiliones. Kunchong-xuebao: jikan = Acta entomologica Sinica / Zhongguo Kunchong Xuehui zhuban. Kunchong-xuebao Bianji Weiyuanhui bianji. - Beijing: Kexue Chubanshe, 3, 503–512.


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